Lawes, the athletic Northampton lock, takes over in the second row from veteran Simon Shaw, who played last week as if the years are beginning to catch up on him.

Both Youngs and Lawes will bring a level of dynamism to the England side after they paid the price for a lack of pace and penetration in the first Test.

Much to the surprise of the Australians, Johnson has resisted the temptation to bring Jonny Wilkinson back into the team and stuck by fly-half Toby Flood and inside centre Shontayne Hape.

Mike Tindall has recovered from a dead leg to be named at outside centre but there is no place in the 22-man squad for Mathew Tait, whose seat on the bench has been taken by Delon Armitage.

For Australia, Matt Giteau and Will Genia will both start.

Wallabies head coach Robbie Deans has decided to restore Genia at scrum-half, despite Luke Burgess’ stand-out performance in Australia’s 27-17 first-Test victory.

Genia has overcome a knee ligament strain while Giteau is back at inside centre in place of Berrick Barnes after recovering from a hip problem which forced him to miss last weekend’s game.

Deans has left Australia’s tight five unchanged, keeping faith with the inexperienced front row that were on the receiving end of a hammering from the England pack in Perth last weekend.

The Wallabies had hoped to recall Tatafu Polota-Nau but the hooker aggravated an ankle problem while playing for the Australian Barbarians against England on Tuesday and he will now undergo surgery.

Queensland prop James Slipper will once again cover both tighthead and loosehead prop off the bench, where he is joined by Adam Ashley-Cooper after the versatile full-back shook off an ankle injury.